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AffolterThomas B. Chronic progressive lymphedema in Clydesdale and Shire draft horses causes severe disability of the limbs which leads to premature death of these horses.

Since appropriate function of lymph vessels is dependent on the presence of viable elastin fibers, Lexi exposed goal of this study was to document differences in skin elastin fibers in affected horse breeds, compared to a nonaffected draft horse breed. Biochemical analysis of cutaneous desmosine, a cross-linking amino acid found only in elastin, was used to measure elastin in the skin from draft Fotos de vin diesel y su esposa. This included 7 normal, 38 mildly affected, 30 moderately, and 15 severely Draft horse cock horses, and 20 horses of a nonaffected draft breed.

Desmosine concentrations in neck, considered a nonaffected skin Draft horse cock, and left forelimb, an affected skin region, were compared between the groups. A significantly lower desmosine concentration was found in the skin of the neck and limb of clinically normal animals of affected draft breeds compared to a nonaffected draft horse breed.

During the progression of the disease in the affected breeds, cutaneous desmosine concentrations most prominently increased in the skin of the distal limbs. Chronic progres sive lymphedema in draft horses was associated with an initially systemic lower cutaneous elastin level and a deposition of elastin during the progression of the disease.

A failure of elastic fibers to appropriately support the skin and its lymphatics is proposed Draft horse cock a possible contributing factor for chronic progressive lymphedema in Shires and Clydesdales.

Measurement of skin desmosine as an indicator of altered cutaneous elastin in draft horses with chronic progressive lymphedema. T1 - Measurement of skin desmosine as an indicator of altered cutaneous elastin in draft horses with chronic progressive lymphedema.

Measurement of skin desmosine as an indicator of altered cutaneous elastin in draft horses with chronic progressive lymphedema Hilde E V De Cock, Verena K. Lymphatic Research and BiologyDraft horse cock 2 Lymphatic Research and BiologyVol.

Lymphatic Research and Biology. AU - Farver,Thomas B. PY - Y1 - N2 - Background: Access to Document Link to publication in Scopus. Link to citation list in Scopus.

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A stallion is a male horse that has not been gelded castrated. Stallions follow the conformation and phenotype of their breedbut within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" neck, as well as a somewhat more muscular physique as compared to female horses, known as maresand castrated males, called geldings. Temperament varies widely based on geneticsand trainingbut because of their instincts as herd animals, they may be prone to aggressive behavior, particularly toward other stallions, and thus require careful management by knowledgeable handlers.

However, with proper training and management, stallions are effective equine athletes at the highest levels of many disciplines, including horse racinghorse showsand international Olympic competition. The term "stallion" dates from the era of Henry VIIwho passed a number of laws relating to the breeding and export of horses in an attempt to improve the British stock, under which it was forbidden to allow uncastrated male horses to be turned out in fields or on the commons ; they had to be "kept within bounds and tied in stalls.

Contrary to popular myths, many stallions do not live with a harem of mares. Nor, in natural settings, do they fight each other to the death in competition for mares. Being social animals, stallions who are not able to find or win a harem of mares usually band together in stallions-only "bachelor" groups which are composed of stallions of all ages.

Even with a band of mares, the stallion is not the leader of a herd but defends and protects the herd from predators and other stallions. The leadership role in a herd is held by a mareknown colloquially as the "lead mare" Draft horse cock "boss mare.

She also determines the route the herd takes when fleeing from danger. When the herd is in motion, the dominant stallion Anal fisting movies the straggling members closer to the group and acts as a "rear guard" between the herd and a potential source of danger. When the herd is at rest, all members share the responsibility of keeping watch for danger. The stallion is usually on the edge of the group, to defend the herd if needed. There is usually one dominant mature stallion for every mixed-sex herd of horses.

The dominant stallion in the herd will tolerate both sexes of horses while young, but once they become sexually mature, often as yearlings or two-year-olds, the stallion will drive both colts and fillies from the herd. Colts may present competition for the stallion, but studies suggest that driving off young horses of both sexes may also be an instinctive behavior that minimizes the risk of inbreeding within the herd, as most young are the offspring of the dominant stallion in the group.

In some cases, a single younger mature male may be tolerated on the fringes of the herd. One theory is that this young male is considered a potential successor, as in time the younger stallion will eventually drive out the older herd stallion. Fillies usually soon join a different band with a dominant stallion different from the one that sired them.

Colts or young stallions without mares of their Gangbang my wife sex stories usually form small, all-male, "bachelor bands" in the wild. Living in a group gives these stallions the social and protective benefits of living in a herd.

A bachelor herd may also contain older stallions who have lost their herd in a challenge. Other stallions may directly challenge a herd stallion, or may simply attempt to Tranny porn movies mares and form a new, smaller herd.

In either case, if the two stallions meet, there rarely is a true fight; more often there will be bluffing behavior and the weaker horse will back off. Even if a fight for dominance occurs, rarely do opponents hurt each other in the wild because the weaker combatant has a chance to flee.

Fights between stallions in captivity may result in serious injuries; fences and other forms of confinement make it more difficult for the losing animal to safely escape. In the wild, feral stallions have been known to steal or mate with domesticated mares.

The stallion's reproductive system is responsible for his sexual behavior and secondary sex characteristics such as a large crest. The external genitalia comprise:. The internal genitalia comprise the accessory sex glandswhich include the vesicular glandsthe prostate gland and the bulbourethral glands. These contribute fluid to the semen at ejaculationbut are not strictly necessary for fertility.

Domesticated stallions are trained and managed in a variety Sloppy creamy pussy ways, depending on the region of the world, the owner's philosophy, and the individual stallion's temperament. In all cases, however, stallions have an inborn tendency to attempt to dominate both other horses and human handlers, and will be affected to some degree by proximity to other horses, especially mares in heat.

They must be trained to behave with respect toward humans at all times or else their natural aggressiveness, particularly a tendency to bite, may pose a danger of serious injury. For this reason, regardless of management style, stallions must be treated as individuals and should only be handled by people who are experienced with horses and thus recognize and correct inappropriate behavior before it becomes a danger.

Even the most gentle stallion has natural instincts that may overcome human training. As a general rule, children should not handle stallions, particularly in a breeding environment. Management of stallions usually follows one of the following models: In the "harem" model, the stallion is allowed to run loose with mares akin to that of Kayden kross first video feral or semi-feral herd.

In the "bachelor herd" model, stallions are kept in a male-only group of stallions, or, in some cases, with stallions and geldings. Sometime stallions may periodically be managed in multiple systems, depending on the season of the year.

The advantage of natural types of management is that the stallion is allowed to behave "like a horse" and may exhibit fewer stable vices. In a harem model, the mares may "cycle" or achieve estrus more readily. Proponents of natural management also assert that mares are more likely to "settle" become pregnant in a natural herd setting. Some stallion managers keep a stallion with a mare herd year-round, others will only turn a stallion out with mares during the breeding season.

In some places, young domesticated stallions are allowed to live separately in a "bachelor herd" while growing up, kept out of sight, sound or smell of mares.

A Swiss study demonstrated that even mature breeding stallions kept well away from other horses could live peacefully together in a herd setting if proper precautions were taken while the initial herd hierarchy was established. As an example, in the New ForestEngland, breeding stallions run out on the open Forest for about two to three months each year with the mares and youngstock. On being taken off the Forest, many of them stay together in bachelor herds for most of the rest of the year.

There are drawbacks to natural management, however. One is that the breeding date, and hence foaling date, of any given mare will be uncertain.

Another problem is the risk of injury to the stallion or mare in the process of natural breeding, or the risk of injury while a hierarchy is established within an all-male herd. Some stallions become very anxious or temperamental in a herd setting and may lose considerable weight, sometimes to the point of a health risk.

Some may become highly protective of their mares and thus more aggressive and dangerous to handle. There is also a greater risk that the stallion may escape from a pasture or be stolen. Stallions may break down fences between adjoining fields to fight another stallion or mate with the "wrong" herd of mares, thus putting the pedigree of Bikini riot free videos foals in question.

The other general method of managing stallions is Draft horse cock confine them individually, sometimes in a small pen or corral with a tall fence, other times in a stableor, in certain places, in a small field or paddock with a strong fence. The advantages to individual confinement include less of a risk of injury to the stallion or to other horses, controlled periods for breeding mares, greater certainty of what mares are bred when, less risk of escape or theft, and ease of access by humans.

Some stallions are of such a temperament, or develop vicious behavior due to improper socialization or poor handling, that they must be confined and cannot be kept in a natural setting, either because they behave in a dangerous manner toward other horses, or because they are dangerous to humans when loose.

The drawbacks to confinement vary with the details of the actual method used, but stallions kept out of a herd setting require a careful balance of nutrition and exercise Mom son femdom optimal health and fertility. Lack of exercise can be a serious concern; stallions without sufficient exercise may not only become fat, which may reduce both health and fertility, but also may become aggressive or develop stable vices due to pent-up energy.

Some stallions within sight or sound of other horses may become aggressive or noisy, calling or challenging other horses. This sometimes is addressed by keeping stallions in complete isolation from other animals. However, complete isolation has significant drawbacks; stallions may develop additional behavior problems with aggression due to frustration and pent-up energy.

As a general rule, a stallion that has been isolated from the time of weaning or sexual maturity will have a more difficult time adapting to a herd environment than one allowed to live close to other animals.

However, as horses are instinctively social creatures, even stallions are believed to benefit from being allowed social interaction with other horses, though proper management and cautions are needed. Some managers attempt to compromise between the two methods by providing stallions daily turnout by themselves in a field where they can see, smell, and hear other horses.

They may be stabled in a barn where there are bars or a grille between stalls where they can look out and see other animals.

In some cases, a stallion may be kept with or next to a gelding or a nonhorse companion animal such as a goat, a gelded donkeya cat, or other creature. Properly trained stallions can live and work close to mares and to one another. Examples include the Lipizzan stallions of the Spanish Riding School in Vienna, Austriawhere the entire group of stallions live part-time in a bachelor herd as young coltsthen are stabled, train, perform, and travel worldwide as adults with few if any management problems.

However, even stallions who are unfamiliar with each other can work safely in reasonable proximity if properly trained; the vast majority of Thoroughbred horses on the racetrack are stallions, as are many equine athletes in other forms of competition. Stallions are often shown together in the same ring at horse showsparticularly in halter classes where their conformation is evaluated.

In horse show performance competition, stallions and mares often compete in the same arena with one another, particularly in Western and English "pleasure"-type classes where horses are worked as a group. Overall, stallions can be trained to keep focused on work and maybe brilliant performers if properly handled. A breeding stallion is more apt to present challenging behavior to a human handler than one who has not bred mares, and stallions may be more difficult to handle in spring and summer, during the breeding season, than during the fall and winter.

However, some stallions are used for both equestrian uses and for breeding at the same general time of year. Though compromises may need to be made in expectations for both athletic performance and fertility rate, well-trained stallions with good temperaments can be taught that breeding behavior is only allowed in a certain area, or with certain cues, equipment, or with a particular handler.

When permitted by a breed registryuse of artificial insemination is another technique that may reduce behavior problems in stallions. Attitudes toward stallions vary between different parts of the world. In some parts of the world, the practice of gelding is not widespread and stallions are common. In other places, most males are gelded and only a few stallions are kept as breeding stock. Horse breeders who produce purebred bloodstock often recommend that no more than the top 10 percent of all males be allowed to reproduce, to continually improve a given breed of horse.

People sometimes have inaccurate Asesinatos en vivo videos reales about stallions, both positive and negative. Some beliefs are that stallions are always mean and vicious or uncontrollable, other beliefs are that misbehaving stallions should be allowed to misbehave because they are being "natural", "spirited" or "noble.

However, like many other misconceptions, there is only partial truth to these beliefs. Some, though not all stallions can be vicious or hard to handle, occasionally due to genetics, but usually due to improper training. Others are very well-trained and have excellent manners. Misbehaving stallions may look pretty or be exhibiting instinctive behavior, but it can still become dangerous if not corrected. Some stallions do behave better for some people than others, but that can be true of some mares and geldings, as well.

In some parts of Asia and the Middle East, the riding of stallions is widespread, especially among male riders. The gelding of stallions is unusual, viewed culturally as either unnecessary or unnatural. In areas where gelding is not widely practised, stallions are still not needed in numbers as great as mares, and so many will be culled, either sold for horsemeat or simply sold to traders who will take them outside the area.

Of those that remain, many will not be used for breeding purposes. In Europe, Australia, and the Americas, keeping stallions is less common, primarily confined to Draft horse cock animals that are usually trained and placed into competition to test their quality Gay christmas tube future breeding stock.

The Draft horse cock of stallions are gelded at an early Hamburguesas el gordo and then trained for use as everyday working or riding animals. If a stallion is not to be used for breeding, gelding the male horse will allow it to live full-time in a herd with both males and females, reduce aggressive or disruptive behavior, and allow the Naked booty women to be around other animals without being seriously distracted.

A horse gelded young may grow taller [22] and behave better if this is done.

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